SUN Chong-bo~(1,2) JIANG Gui-hua~1 SHI Ji-sen~2 ZHANG Hui-qin~1 WU Yan-jun~1 GUO Fang-qi~1 XIE Ming~1(1.Institute of Horticulture,Zhejiang Academy of Agricultural Sciences,Hangzhou,Zhejiang 310021;2.Key Laboratory of Forest Tree Genetics and Gene Engineering of State Forestry Administration,Nanjing Forestry University,Nanjing,Jiangsu 210037)